If an app is developed using Storyboard in Xcode 4.2, can this app run in iOS 4.x devices?
No, it can't. Storyboards are only available in iOS 5.
From the UIStoryboard class reference:
Available in iOS 5.0 and later
From Converting to Storyboards Release Notes:
Storyboarding is a new way to create user interfaces for iOS applications, beginning with iOS 5 and Xcode 4.2.
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