I would like to know whether there are any websites (not toy projects) that have been successfully built on top of GAE (excluding google) using whatever programming languages or frameworks that are available.
Yes, of course there are. We've featured a few of them on the blog in the past, including webfilings, Wolfire games, and Neptune's Pride, to name a few.
Overgrowth looks fairly big.
The App Engine gallery, but it has been closed.
And there is also StackPrinter...
See: 5 cool web applications hosted on Google App Engine
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