Can I change readonly_fields in my TranslationAdmin class dependent on the value of a certain field in the Translation being viewed? If so, how would I do that?
The only thing I've come up with is to make a widget that looks at the Translation and decides whether to be a readonly widget or not, but that seems like overkill for this. 
I think you have misunderstood what fieldsets are. It's just a way for you to be able to group the fields on a Model Admin Page.
One of the most powerful parts of Django is the automatic admin interface. It reads metadata from your models to provide a quick, model-centric interface where trusted users can manage content on your site. The admin's recommended use is limited to an organization's internal management tool.
The attribute prepopulated_fields tells the admin application to automatically fill the field slug - in this case with the text entered into the name field.
You can inherit the function get_readonly_fields() in your admin and set readonly fields according your model's certain field value
 class TranslationAdmin(admin.ModelAdmin):
        ...
        def get_readonly_fields(self, request, obj=None):
            if obj.certainfield == something:
                return ('field1', 'field2')
            else:
                return super(TranslationAdmin, self).get_readonly_fields(request, obj)
I hope it will help you.
Here is an example of:
Inheriting from that ModelAdmin (ShapefileSetAdmin) and adding an additional value to readonly_fields
class GISDataFileAdmin(admin.ModelAdmin):
    # Note(!): this is a list, NOT a tuple
    readonly_fields = ['modified', 'created', 'md5',]
class ShapefileSetAdmin(GISDataFileAdmin):    
    def get_readonly_fields(self, request, obj=None):
        # inherits readonly_fields from GISDataFileAdmin and adds another
        # retrieve current readonly fields 
        ro_fields = super(ShapefileSetAdmin, self).get_readonly_fields(request, obj)
        # check if new field already exists, if not, add it
        #
        # Note: removing the 'if not' check will add the new read-only field
        #   each time you go to the 'add' page in the admin
        #   e.g., you can end up with:
        # ['modified', 'created', 'md5', 'shapefile_load_path', 'shapefile_load_path, 'shapefile_load_path', etc.]
        #
        if not 'shapefile_load_path' in ro_fields:
            ro_fields.append('shapefile_load_path')
        return ro_fields
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